Unity4 DXT APKs don't support S3TC, can't publish to Play Store

(I've taken over a long-running android unity project after the previous developer left (don't they always?) so there may be some misisng holes here in my knowledge)

After building all four android subtargets through unity 4 and uploading them to the google play store developer console, the new v2 console is telling me that I can't publish because of the following reasons:

 This configuration cannot be published for the following reason(s):
     It is forbidden to downgrade devices which previously matched version 2009 to version 1057, which would occur when 
     API levels in range 8+ and 
     Screen layouts containing any of [small, normal, large, xlarge] and 
     Native platforms containing any of [armeabi-v7a] and 
     OpenGL ES versions in range 2.0+ and 
     OpenGL textures containing any of [GL_EXT_texture_compression_s3tc] and containing any of [GL_OES_compressed_ETC1_RGB8_texture] but excluding all of [GL_AMD_compressed_ATC_texture, GL_ATI_texture_compression_atitc, GL_EXT_texture_compression_dxt1, GL_EXT_texture_compression_dxt5, GL_IMG_texture_compression_pvrtc] and 
     Features containing all of [android.hardware.sensor.ACCELEROMETER, android.hardware.TOUCHSCREEN, android.hardware.touchscreen.MULTITOUCH, android.hardware.WIFI].

I am indeed making an update at version *057, but the previous version was *055, not *009.

I've dug out version 2009, and it indeed supports S3TC. No version after that does including my new update.

Was S3TC support removed from Unity at some point? How can I convince the v2 developer console to let me publish? Is there any more information I can provide you with?